Professor
Biology
Cohen Science Center 211
michael.steele@wilkes.edu
(570) 408-4763
Michael A. Steele
Professor of Biology and H. Fenner Chair of Research Biology
Ph.D. 1988. Wake Forest University
Research Interests:
Evolutionary ecology; behavioral, population and community ecology of vertebrates;
plant-herbivore interactions, foraging behavior, seed predation, and seed dispersal
Honors and Awards:
2004 Innovative Teaching Award, (student nomination), Wilkes University
2003 Outstanding Teaching/ Research Award, Wilkes University (top 5% of faculty each
year)
2002 Outstanding Teaching/ Research Award, Wilkes University
2001 Outstanding Teaching/ Research Award, Wilkes University
1997-98 Fulbright Fellowship to Mexico
1997 Outstanding Teaching/ Research Award, Wilkes University
1996 Outstanding Teaching/Research Award, Wilkes University
1994 Millersville University, Biology Department, Alumni Fellow Research Award
1994 Wake Forest University, Biology Department, Alumni Seminar Award
1994-95 Sigma Xi, President, Wilkes University Chapter
1994 Outstanding Teaching/Research Award, Wilkes University
1993 January, 1993 Commencement Address, Wilkes University
1992 Carpenter Outstanding Teaching Award (one-time only, university teaching award
for top faculty member among 153 faculty; selected by faculty committee)
1991 Outstanding Teaching/Research Award, Wilkes University
1990 Full Member, Sigma Xi Honor Society
1988 Sigma Xi Outstanding Graduate Award in Biological Sciences
1988 Outstanding Graduate Research Award in Biological Sciences, Benkman Scientific/
WFU Graduate Student Association
1985 Associate Member Sigma Xi Honor Society
1982 Phi Kappa Phi Honor Society
1982 Outstanding Senior Biology Award (in dept. of 450 majors), The Assoc. of State
1982 College and University Biology Faculty
Michael A. Steele
Permanent Address:
Department of Biology
Wilkes University
Wilkes-Barre, PA 18766
Education:
Postdoctoral Research Associate, N. C. State University, Raleigh, N.C.
Wake Forest University, Ph.D. Biology (Ecology and Behavior) May, 1988
Millersville University of Pennsylvania, B.S. Biology (High Honors/Magna Cum Laude),
1982
Teaching Experience and Positions:
2005-P H. Fenner Chair of Research Biology (Endowed Research Chair), Wilkes University
2003-P Professor of Biology, Wilkes University, (Primary courses: Animal Behavior,
Parasitology, Ecology, Evolution, Evol. Ecology, Plant-Animal Interactions)
1996-03 Associate Professor of Biology, Wilkes University
1989-95 Assistant Professor of Biology, Wilkes University
1988 Visiting Lecturer in Biology (Zoology), Salem College, Winston-Salem, NC
1986-87 Teaching Assistantship, Wake Forest University (General Biology, Organismal
Biology, Ecology)
1985-86 Co-director of a graduate study group in Evolution Ecology, Wake Forest University
1985-89 Occasional lectures in Vertebrate Zoology and Ecology, Wake Forest University
1982-83 Teaching Assistantship, Wake Forest University
1979-82 Undergraduate Teaching Assistant and Tutor, Millersville University of PA.
Research Experience and Positions:
2005-P Adjunct Associate Professor, Dept. of Forestry and Natural Resources, Purdue
University
2005-P Research Associate, Hawk Mountain Sanctuary
1997-98 Fulbright Fellow, UNAM, Mexico City and BUAP, Puebla Mexico
1992-P Research Associate, Powdermill Biological Station, Carnegie Museum of Natural
History
1988-89 Postdoctoral Research Associate, North Carolina State Univ., Raleigh, N.C.
1985-88 Research Associate, Wake Forest University
1983-85 Student Curator of Mammal Collections, Wake Forest University
1983-85 Research Assistantship, Wake Forest University
Awards:
2004 Innovative Teaching Award, (student nomination), Wilkes University
2003 Outstanding Teaching/ Research Award, Wilkes University (top 5% of faculty each
year)
2002 Outstanding Teaching/ Research Award, Wilkes University
2001 Outstanding Teaching/ Research Award, Wilkes University
1997-98 Fulbright Fellowship to Mexico
1997 Outstanding Teaching/ Research Award, Wilkes University
1996 Outstanding Teaching/Research Award, Wilkes University
1994 Millersville University, Biology Department, Alumni Fellow Research Award
1994 Wake Forest University, Biology Department, Alumni Seminar Award
1994-95 Sigma Xi, President, Wilkes University Chapter
1994 Outstanding Teaching/Research Award, Wilkes University
1993 January, 1993 Commencement Address, Wilkes University
1992 Carpenter Outstanding Teaching Award (one-time only, university teaching award
for
top faculty member among 153 faculty; selected by faculty committee)
1991 Outstanding Teaching/Research Award, Wilkes University
1990 Full Member, Sigma Xi Honor Society
1988 Sigma Xi Outstanding Graduate Award in Biological Sciences
1988 Outstanding Graduate Research Award in Biological Sciences, Benkman Scientific/
WFU Graduate Student Association
1985 Associate Member Sigma Xi Honor Society
1982 Phi Kappa Phi Honor Society
1982 Outstanding Senior Biology Award (in dept. of 450 majors), The Assoc. of State
1982 College and University Biology Faculty
Research Support: (Total support > $2.5 million; recent support in the last 5 yrs
includes 10 grants for > $1.7 million) Student collaborators in bold.
Recent and Current Grants:
1999-04 National Science Foundation: “A multidisciplinary approach to understanding
interactions between food-hoarding animals and the oaks” (Steele PI, $833,000.00,
completed in 04)
2001-04 Pennsylvania Game Commission and USFWS: “A state-wide monitoring program for
the threatened Northern Flying Squirrel” (Steele and Mahan, $107,000.00, completed
in 04)
2002-05 American Association for the Advancement of Science (AAAS) and MERK Foundation.
Biology and Chemistry Undergraduate Research Program (Terzaghi et al. $60,000.00)
2003-05 Pennsylvania Game Commission and USFWS: “Rare threatened, and endangered vertebrates
of PA: A guide for research, management, and conservation” (Steele and Merritt, $40,000.00)
2003-05 Pennsylvania Game Commission and USFWS: “Continuation of a state-wide monitoring
program for the threatened Northern Flying Squirrel” (Steele, Mahan and Turner, $35,000.00)
2005-06 Pennsylvania Wild Resource Conservation Fund: “Long-term monitoring of small
mammals in Pennsylvania” (Steele and Zegers, $42,000.00)
2005-06 Pennsylvania Wild Resource Conservation Fund: “Forest stand age, composition,
and structure associated with the State Endangered Northern Flying Squirrel” (Mahan
et al. $25,000.00)
2005-07 Pennsylvania Game Commission and USFWS: “Eastern hemlock and mixed coniferous
forested ecosystems: “distribution and use by the endangered northern flying squirrel”(Mahan
and Steele, $65,083.00)
2004-05 Wilkes University Faculty Development Grant: “Agro farming by tree squirrels”
(Steele, McEuen, and Miller, $3000.00)
2007-2010 National Science Foundation: “Collaborative Research: Effects of landscape
structure on animal-mediated acorn dispersal and seedling recruitment in the Central
Hardwoods.” (Swihart [Purdue] and Steele CoPIs, $475,000.00)
2007-2009 Pennsylvania Game Commission and USFWS: Final publications costs for “Rare
threatened, and endangered vertebrates of PA: A guide for research, management, and
conservation” (Steele, Brittingham, Maret, and Merritt, $38,000.00)
Additional Fund-Raising:
2003-05 Coordinated effort with Development Office of Wilkes University to raise $250,000
for establishment of Turoczi Genomics Facility
Additional Research Support:
2001-02 PA Dept of Education ($280,000)
2001 Wilkes University Faculty Development ($1500)
1997 Fulbright Fellowship ($48k)
1997 Wilkes University Faculty Development ($2,000)
1995-97 Pennsylvania Wild Resource Conservation Fund ($32,000)
1995 U.S. Fish and Wildlife Service Grant ($2500)
1994-96 National Science Foundation (ROA) Grant (with J. Wolff, $17,000)
1994 Wilkes University Faculty Development ($1,500)
1993 Pennsylvania Wild Resource Conservation Fund ($6,000)
1992 Wilkes University Faculty Development Grant ($1,950)
1992 Wilkes University Faculty Development Grant ($1,200)
1990-91 Pennsylvania Wild Resource Conservation Fund ($34,100)
1990 Wilkes University Faculty Development Grant ($2,450)
1990 Highlands Biological Station Research Grant ($600)
1990 North Carolina Non-Game Research Grant ($3,000)
1900 North Carolina Non-Game Research Grant ($2,000)
1989 Highlands Biological Station Postdoctoral Research Grant ($2,000)
1987 Wake Forest University Summer Research Fellowship ($800)
1986 American Society of Mammalogists Research Grant ($500)
1986 North Carolina Wildlife Federation Scholarship and Research Grant ($500)
1985 Sigma Xi Grant-in-Aid of Research (National Chapter) ($250)
1983-87 Wake Forest University Travel Grants for paper presentations ($1,200)
1983 Wake Forest University Summer Research Fellowship
1981 Millersville University Alumni Soc. Research Grant ($350)
Research Interests:
Evolutionary ecology; behavioral, population and community ecology of vertebrates;
plant-herbivore interactions, foraging behavior, seed predation, and seed dispersal
Professional Societies:
American Society of Mammalogists
Animal Behaviour Society
American Association for the Advancement of Science
Ecological Society of America
Sigma Xi Honor Society
American Society of Naturalists
Pennsylvania Academy of Science
International Society of Behavioral Ecologists
The American Chestnut Foundation
Professional Reviewer:
Behaviour
Behavioural Processes
Animal Cognition
Journal of Zoology, London
Journal of Functional Ecology
Canadian Journal of Botany
Acta Oecologia
Acta Theriologica
Australian Journal of Zoology
Forest Ecology and Management
Oikos
Behavioral Ecology
Modrona (Plant Journal)
Eco-science
Western North American Naturalist
Southwestern Naturalist
Northeastern Naturalist
Ecology
Ecological Monographs
Journal of Wildlife Management
Journal of Mammalogy
Pennsylvania Academy of Science
American Midland Naturalist
U.S. Fish and Wildlife Service
Mammalian Species
Wildlife Society Bulletin
Natural History
Various Symposia Volumes
Johns Hopkins University Press
University of New Mexico Press
National Science Foundation
U.S. Department of Agriculture
U.S. Geological Service
Fish and Wildlife Service
National Geographic Society
Various Textbooks
Various Edited Volumes
Other Professional Activities:
Convener: International Theriological Congress Symposium: Ecological and Evolutionary
Aspects of Plant-mammal Interactions (1997)
Convener: International Colloquium on Ecology of Tree Squirrels (1994)
Pennsylvania Biological Survey Committee (State Mammal Technical Committee, 1991-P)
LACAWAC Sanctuary: Scientific Advisory Board (1993-P)
American Society of Mammalogists: Program Committee (1993-1996), Honors / Grant Committee
(2001-P)
U.S. Fish and Wildlife Service Endangered Species Recovery Team (Delmarva Fox Squirrel
(1991-P)
University Service (abbreviated activities):
Chair of Division of Biology, Chemistry and Health Science (2002-2005)
Acting Chair of Biology, Chemistry and Health Science (2001)
Core Curriculum Development Committee
Assistant to the Dean on Research Activities in School of Science and Engineering
(1994-97)
Teacher Recognition and Enhancement Committee (1994-97)
Co-Chair, AIDS Awareness Committee (1992-95)
Chair, Animal Care and Use Committee (1990-95)
Education Standards Committee (1991-92)
Graduate Studies Committee (1992-95)
Representative of the School of Science and Engineering to University Strategic Long-range
Planning Committee (1991)
President, Sigma Xi, Wilkes University Chapter (1994-96)
Co-Advisor to Biology, Premedical, and Student AIDS Awareness Organizations (1990-95)
Invited Research Seminars and Keynote addresses:
2006 Central Connecticut State University, New Britain, CT
2006 Fourth International Colloquium on Tree Squirrels, Plenary Lecture, India
2005 International Mammal Congress, Workshop on Tree squirrels, Sapporo, Japan
2005 International Mammal Congress (Invited Symposium Presentation, Symposium on Mammal
Plant Interactions), Sapporo, Japan
2005 Fourth international Symposium on Frugivory and Seed Dispersal (Plenary Lecture),
Brisbane Australia
2004 Millersville University, Millersville, PA
2004 Pennsylvania State University, State College, PA
2004 Fordham, University, New York City
2004 Miami University of Ohio, Oxford, OH
2003 University of Maryland, Environmental Research Laboratory, Frostburg, MD.
2003 American Chestnut Foundation, PSU, State College PA
2003 Third International Colloquium on Tree Squirrels, Plenary Lecture, Northumberland
, UK
2002 Scranton University, Scranton, PA
2002 Pymentuning Center for Ecology, University of Pittsburgh
2002 Northern Arizona University, Flagstaff, AZ
2002 University of Arizona, Tucson, AZ
2002 Keynote Address, Upland Oak Symposium, University of Arkansas, Feyetteville,
AR
2001 Millersville University, Millersville PA
2000 Wake Forest University, Winston Salem, NC
2000 Purdue University, Department of Forest Resources, Lafayette, IN
2000 Lycoming College, Williamsport, PA
1999 Franklin and Marshal College, Department of Biology, Lancaster, PA
1998 Pennsylvania State University, Altoona, PA
1998 Fordham University, New York City
1998 Centro de Ecologia, UNAM, Mexico City, Mexico
1997 Africam Zoological Park, Puebla, Mexico
1997 Benemerita Universidad de Puebla, Puebla, Mexico
1997 International Theriological Congress, (Invited Symposium Presentation, Symposium
on Mammal Plant Interactions), Acapulco, Mexico
1997 Juniata University, Juniata, PA
1996 LACAWAC Sanctuary (Lehigh University Summer REU Research Program)
1996 Scranton University, Scranton
1995 Villanova University, Philadelphia
1994 Wake Forest University, Winston-Salem, NC
1993 Millersville University of Pennsylvania, Millersville, PA
1993 Bucknell University, Lewisburg, PA
1993 King’s College, Wilkes-Barre, PA
1992 Millersville University of Pennsylvania, Millersville, PA
1991 Pennsylvania State University, State College, PA
1991 East Stroudsburg University, East Stroudsburg, PA
1990 Clemson University, Clemson, SC
1989 Wilkes University, Wilkes-Barre, PA
1989 Salem College, Winston-Salem, NC
1988 Millersville University of Pennsylvania, Millersville, PA
1986 Sigma Xi Research Award Presentation, Bowman Gray School of Medicine,
1986 Wake Forest University, Winston-Salem, NC
Refereed Books:
Steele, M. A. and J. L. Koprowski. 2001. North American Tree Squirrels. Smithsonian Institution Press. (Reprinted in Spring of 2002).
Steele, M. A., J. Merritt, and D. A. Zegers, eds. 1998 The Evolution and Ecology of Tree Squirrels. Virginia Museum of Natural History, Special Publication 6., Martinsville, VA.
M. Steele, M. Brittingham, T. Maret, and J. Merritt (eds.) (In Review). Terrestrial Vertebrates of Concern In Pennsylvania: A Guide for Research, Conservation and Management.
Refereed Papers:
Steele, M. A, S. L. Halkins, P. D. Smallwood, T. McKenna, K. Mistopolus, and M. Beam,. In review. Cache protection strategies of a scatter-hoarding rodent: Do tree squirrels engage in behavioral deception? Animal Behaviour.
Koprowski, J. L. and M. A. Steele. In review. Tamisciurus mearnsii Mammalian Species.
Steele, M A. and D. A. Zegers The Delmarava fox squirrel (Sciurus niger vulpinus). In review M. Steele, M. Brittingham, T. Maret, and J. Merritt (eds.). Terrestrial Vertebrates of Pennsylvania: Species of Conservation Concern.
Steele, M A. The Eastern fox squirrel (Sciurus niger vulpinus). In press. In M. Steele, M. Brittingham, T. Maret, and J. Merritt (eds.). Terrestrial Vertebrates of Pennsylvania: Species of Conservation Concern.
Steele, M.A., G. Turner, C Mahan. The northern flying squirrel (Glaucomys sabrinus). In press. In M. Steele, M. Brittingham, T. Maret, and J. Merritt (eds.) (In review Terrestrial Vertebrates of Pennsylvania: Species of Conservation Concern.
Hansen, D., S. Vigallon, J. Wilson, L. Hamilton, M. Steele, R. Innes, S. Krause, M. Varner, B. Woodbridge, and T. George. In press. Ecology and management of goshawk foraging areas. In: Woodbridge, B., J. Keane, D. Hansen, and J. Dunk (tech. eds.). The northern goshawk in California: a technical assessment of its current status and ecology. Gen. Tech. Report, PSW-GTR-xxx, USDA Forest Service.
Steele, M. A. In press. Evolutionary interactions between tree squirrels and trees: A review and synthesis. Current Science.
Moore, J. E., A. B. McEuen, R. K. Swihart, T. A. Contreras, and M. A. Steele. In press. Determinants of seed-removal distance by scatter-hoarding rodents in deciduous forests. Ecology.
Steele, M. AJ. E. Carlson, P.D. Smallwood, A. B. McEuen, T. A. Contreras, and W. B. Terzaghi. 2007. Linking seed and seed shadows: A case study in the oaks (Quercus) In A. J. Dennis, E. W. Schupp, R. J. Green, D. A. Wescott (eds). Seed Dispersal: Theory and its Application in a Changing World. CAB International.
Steele, M. A. 2007 Biology of the Sciuridae: A global perspective. Review of Squirrels: The Animal Answer Guide. Johns Hopkins University Press. Journal of Mammalian Evolution, DOI 10.1007/210914-007-9047-5.
Steele, M. A., S. Manierre, T. Genna, T. Contreras, P. D. Smallwood, and M. Pereira. 2006. The innate basis of food hoarding decisions in grey squirrels: evidence for behavioral adaptations to the oaks. Animal Behaviour, 71, 155-160.
Steele, M. A., B. C. McCarthy, C. H. Keiffer. 2005. Seed dispersal, seed predation, and the American Chestnut. Journal of the American Chestnut Foundation, XIX, 47-54.
Jaffe, G. D. A. Zegers, M. A. Steele, and J. F. Merritt. 2005. Long term patterns of botfly parasitism in Peromyscus maniculatus, P. leucopus, and Tamias striatus. Journal of Mammalogy.
Steele, M. A. and A. B. McEuen. 2005. Atypical acorns appear to allow escape after apical notching by tree squirrels. American Midland Naturalist, 154:450-458.
Steele, M. A., L. Wauters, and K. Larsen. 2005. Selection, predation, and dispersal of seeds by tree squirrels in temperate and boreal forests: Are tree squirrels keystone granivores? In P. M. Forget, S. Vander Wall. J. Lambert, and P. Hulme. Seed Fate: Predation, Dispersal and Seedling Establishment. CAB International.
Steele, M. A., P. D. Smallwood, W. Terzaghi, J. Carlson, T. Contreras, and A. McEuen. 2004. Oak Dispersal Syndromes: Do red and white oaks exhibit different dispersal strategies? Proceedings of the Upland Oak Symposium, October 2002, Technical Publication of the U.S. Forest Service.
Lavenex, P., M. A. Steele, and L. F. Jacobs. 2003. Lack of seasonal variation in the hippocampus: statistics is not the issue. Animal Behaviour, DOI:10.1006
Ceballos, G., H. Zarza, and M. A. Steele 2003. Marmosa candescent Mammalian Species, 725, 1-4.
Ceballos, G., H. Zarza, and M. A. Steele. 2002. Xenomys nelsoni Mammalian Species, 704, 1-3.
Steele, M. A. and P. D. Smallwood. 2002. Acorn dispersal by birds and mammals. In W. McShea, and W. M. Healy (eds.) Oak Forest Ecosystems: Ecology and Management for Wildlife. Johns Hopkins University Press.
Steele, M. A., G. Turner, P. D. Smallwood.,, J. O. Wolff, and J. Radillo 2001. Cache management by small mammals: experimental evidence for the significance of acorn embryo excision. Journal of Mammalogy, 82:35-42
Smallwood, P. D., M. A. Steele, and S. Faeth. 2001. The ultimate basis of the caching preferences of rodents and the oak-dispersal syndrome: tannins, insects, and seed germination. American Zoologist, 41: 840-851.
Steele, M. A., P. D. Smallwood, A, Spunar and E. Nelsen. 2001. The proximate basis of the oak dispersal syndrome: can small mammals smell seed dormancy? American Zoologist, 41:852-864.
Lavenex, P., M. A. Steele, L. F. Jacobs. 2000. Sex differences but no seasonal variation in the hippocampus of food-caching squirrels: a soteriological study. Journal of Comparative Neurology, 425:152-166.
Lavenex, P., M. A. Steele, L. F. Jacobs. 2000. The seasonal pattern of cell proliferation and neuron number in the dentate gyrus of wild adult eastern gray squirrels. European Journal of Neuroscience, 12: 643-648.
Derge, K.*, and M. A. Steele. 2000. The status of the fox squirrel (Sciurus niger) in Pennsylvania. Journal of the Pennsylvania Academy of Science, 72:43-50.
Mahan, C. G., M. A. Steele, M. J. Patrick, and G. Kirkland, Jr. 2000. Status of the northern flying squirrel (Glaucomys sabrinus) in Pennsylvania. Journal of the Pennsylvania Academy of Science, 73:15-21.
Steele, M. A. 1999 Tamiasciurus douglasi. Mammalian Species , 630, 1-8
Browne, R, M. A. Steele, and P. D. Weigl. 1999. Mountaintops as islands: genetic variation of small mammals in the southern Appalachians. In Appalachian Biogeography (R. Eckerlin, ed.). Virginia Museum of Natural History.
Steele, M. A. and R. Powell. 1999. Biogeography of small mammals in the southern Appalachians: Patterns of local and regional abundance. In Appalachian Biogeography ( R. Eckerlin, , ed.). Virginia Museum of Natural History.
Steele, M. A. Tamiasciurus hudsonicus. 1999. Mammalian Species.
Smallwood, P. D., M. A. Steele, E. Ribbens, and W. F. McShea. 1998. Detecting the effects of seed hoarders on the distribution of tree species: Gray squirrels (Sciurus carolinensis) and oaks (Quercus spp.) as a model system. Pp 211-222 in Steele, M. A., J. Merritt, and D. A. Zegers (eds). 1998. The Ecology of Tree Squirrels. Virginia Museum of Natural History, Special Publication 6., Martinsville, VA.
Weigl, P. D. L. Sherman, M. A. Steele, and D.S. Weaver. 1998. Geographic variation in fox squirrels (Sciurus niger): A consideration of size clines, feeding economics, and body size. Pp. 171-184 in Steele, M. A., J. Merritt, and D. A. Zegers (eds). 1998. The Ecology of Tree Squirrels. Virginia Museum of Natural History, Special Publication 6., Martinsville, VA.
Steele, M. A., K.Gavel, and W. Bachman. 1998. Experimental analysis of responses of gray squirrels to physical and chemical characteristics of acorns: implications for seed dispersal. Pp. 223-232 In Steele, M. A., J. Merritt, and D. A. Zegers (eds). 1998. The Ecology of Tree Squirrels. Virginia Museum of Natural History, Special Publication 6., Martinsville, VA.
Steele, M. A., and J. L. Koprowski. 1998. Closing comments and future directions: research on the ecology and evolutionary biology of tree squirrels. Pp. 309-310 in Steele, M. A., J. Merritt, and D. A. Zegers (eds). 1998. The Ecology of Tree Squirrels. Virginia Museum of Natural History, Special Publication 6., Martinsville, VA.
Steele, M. A., L. Z. Hadj-Chikh, and J. Hazeltine. 1996. Caching and feeding behavior of gray squirrels: responses to weevil infested acorns. J. of Mammalogy, 77: 304-314.
Hadj-Chikh, L. Z., M. A. Steele, and P. D. Smallwood. I996. Caching behavior of gray squirrels: test of the perishability and handling-time hypotheses. Animal Behaviour, 52:941-948.
Steele, M. and G. Kwiecinski. 1994. Mammals of Pennsylvania: a checklist with notes on status, habitat, and behavior. Published by the Pennsylvania Game Commission.
Steele, M. and P. Smallwood. 1994. What are squirrels hiding? Natural History, 10:40-46.
Steele, M., T. Knowles, K. Bridle, and E. Simms. 1993. Tannins and partial consumption of acorns: implications for dispersal of oaks by seed predators. American Midland Naturalist, 130:229-238.
Steele, M. and P. Weigl. 1993. The ecological significance of body size in fox squirrels (Sciurus niger) and gray squirrels (Sciurus carolinensis). In Proceedings of the Second Symposium on Southeastern Fox Squirrels, Sciurus niger (N. D. Moncrief, J. W. Edwards, and P.A. Tappe eds.). Virginia Museum of Natural History Special Publication, 1: 57-69.
(multiple authors). 1993. Delmarva Fox Squirrel Recovery Plan, 2nd Revision. U.S. Fish and Wildlife Service, Northeast Region, 65pp.
Steele, M. A. and P. D. Weigl 1992. Energetics and patch use in the fox squirrel: responses to prey density and patch profitability. American Midland Naturalist, 128: 156-167.
Knowles, T., M. A. Steele, and P. D. Weigl. 1989. Survey of rare and endangered vertebrates along the Blue Ridge Parkway in North Carolina. The National Park Service. 1-172pp.
Weigl, P. D., M.A. Steele, L.J. Sherman, J. Ha, T. Sharpe. 1989. The ecology of the fox squirrel in North Carolina: implications for survival in the southeast. Bulletin of Tall Timbers Research Station, 28:1-93
Weigl, P.D. and M.A. Steele. 1989. Book Review: The Natural History of Tree Squirrels
(By John Gurnell. Facts on File Publ. New York, N.Y., and Oxford U.K. 1989). J. Wildlife
Management, 53:273-274.
Non-refereed Papers/Monographs:
Steele, M. A. C. Mahan, and G. Turner. 2004. A Manual for Long-Term Monitoring and Management of the Northern Flying Squirrel (Glaucmys sabrinus) in Pennsylvania. The Pennsylvania Game Commission and U. S. Fish and Wildlife Services, 137 pp.
Steele, M. A., and P. D. Smallwood. 1996. How to plant an oak forest. Wildflower Magazine (by invitation), Spring, 35.
Steele, M. A.. 1996. The mystery of the half-eaten acorn. Dragonfly (NSF supported magazine for young investigators) (by invitation), April, 28-31.
Steele, M. 1993. Habituation and Sensitization. In Magill’s Survey of Social Science: Psychology (Magill, F.N., ed.). Salem Press, Pasadena, Calif. a
Steele, M. 1993. Ethology. In Magill’s Survey of Social Science: Psychology (Magill, F.N., ed.). Salem Press, Pasadena, Calif. a
Steele, M.A. 1991. Learning and Insight. In McGill's Survey of Science: The Life Sciences (Magill, F.N., ed.). Salem Press.a
Steele, M.A. 1991. The Hardy-Weinberg Law. In McGill's Great Events from History of Science (Magill, F.N., ed.). Salem Press.a
Steele, M. A. 1988. Patch use and foraging ecology of the fox squirrel: tests of theoretical predictions. Ph.D. Dissertation, Wake Forest University, Winston-Salem, N.C. 215p.
Numerous Grant Reports not included
Abstracts from Conferences:
Approximately 150 from 1982-Present
Evolutionary ecology; behavioral, population and community ecology of vertebrates; plant-herbivore interactions, foraging behavior, seed predation, and seed dispersal
2004 Innovative Teaching Award, (student nomination), Wilkes University
2003 Outstanding Teaching/ Research Award, Wilkes University (top 5% of faculty each year)
2002 Outstanding Teaching/ Research Award, Wilkes University
2001 Outstanding Teaching/ Research Award, Wilkes University
1997-98 Fulbright Fellowship to Mexico
1997 Outstanding Teaching/ Research Award, Wilkes University
1996 Outstanding Teaching/Research Award, Wilkes University
1994 Millersville University, Biology Department, Alumni Fellow Research Award
1994 Wake Forest University, Biology Department, Alumni Seminar Award
1994-95 Sigma Xi, President, Wilkes University Chapter
1994 Outstanding Teaching/Research Award, Wilkes University
1993 January, 1993 Commencement Address, Wilkes University
1992 Carpenter Outstanding Teaching Award (one-time only, university teaching award for
top faculty member among 153 faculty; selected by faculty committee)
1991 Outstanding Teaching/Research Award, Wilkes University
1990 Full Member, Sigma Xi Honor Society
1988 Sigma Xi Outstanding Graduate Award in Biological Sciences
1988 Outstanding Graduate Research Award in Biological Sciences, Benkman Scientific/
WFU Graduate Student Association
1985 Associate Member Sigma Xi Honor Society
1982 Phi Kappa Phi Honor Society
1982 Outstanding Senior Biology Award (in dept. of 450 majors), The Assoc. of State
1982 College and University Biology Faculty